The main character of this book is a she. Her name is India Opal Buloni. She is 10 years old and has just moved to Naomi, Florida.
At the start of the book, Opal is sad and lonely. She has no friends in Naomi. She also misses her mother, who left the family when she was a great deal younger. She doesn't remember much about her mother, and that makes her sad.
As the story goes on, Opal learns how to meet people and to come out of her shell. She is helped to do that by Winn-Dixie. He helps her meet people and he also helps her start talking to her dad more. That allows her to find out more about her mom.
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